:root {
  --anatomy-ink: #f7fbfd;
  --anatomy-muted: #98afbb;
  --anatomy-line: rgba(156, 199, 215, 0.16);
  --anatomy-panel: rgba(6, 18, 29, 0.92);
  --anatomy-black: #010305;
  --anatomy-teal: #14f1c4;
  --anatomy-gold: #cfaf70;
}

.anatomy-page {
  background:
    radial-gradient(circle at 78% 4%, rgba(20, 241, 196, 0.075), transparent 29rem),
    radial-gradient(circle at 7% 17%, rgba(31, 162, 255, 0.07), transparent 32rem);
}

.anatomy-page strong {
  color: var(--anatomy-ink);
}

.anatomy-hero {
  min-height: calc(100vh - 74px);
  display: grid;
  grid-template-columns: minmax(0, 0.8fr) minmax(520px, 1.2fr);
  align-items: center;
  gap: clamp(42px, 7vw, 98px);
  padding-block: clamp(72px, 9vw, 128px);
}

.anatomy-hero-copy h1 {
  max-width: 760px;
  margin: 0;
  color: var(--anatomy-ink);
  font-size: clamp(3.8rem, 7.6vw, 8rem);
  font-weight: 640;
  letter-spacing: -0.065em;
  line-height: 0.88;
  text-wrap: balance;
}

.anatomy-hero-lede {
  max-width: 650px;
  margin: 30px 0 0;
  color: var(--anatomy-muted);
  font-size: clamp(1.03rem, 1.55vw, 1.22rem);
  line-height: 1.75;
}

.anatomy-hero-facts {
  display: flex;
  flex-wrap: wrap;
  gap: 9px;
  margin: 34px 0 0;
  padding: 0;
  list-style: none;
}

.anatomy-hero-facts li {
  padding: 8px 12px;
  border: 1px solid var(--anatomy-line);
  border-radius: 999px;
  color: #9db4bf;
  font-size: 0.72rem;
  font-weight: 690;
  letter-spacing: 0.04em;
}

.anatomy-hero-media {
  position: relative;
  min-height: clamp(520px, 66vh, 760px);
  overflow: hidden;
  margin: 0;
  border: 1px solid var(--anatomy-line);
  border-radius: 30px;
  background: #02060a;
  box-shadow: 0 42px 100px rgba(0, 0, 0, 0.42);
}

.anatomy-hero-media::after {
  position: absolute;
  inset: 0;
  background:
    linear-gradient(180deg, transparent 42%, rgba(1, 5, 8, 0.88)),
    linear-gradient(90deg, rgba(1, 5, 8, 0.2), transparent 45%);
  pointer-events: none;
  content: "";
}

.anatomy-hero-media img {
  width: 100%;
  height: 100%;
  min-height: inherit;
  display: block;
  object-fit: cover;
}

.anatomy-hero-media figcaption {
  position: absolute;
  z-index: 1;
  right: 24px;
  bottom: 23px;
  left: 24px;
  display: flex;
  align-items: end;
  justify-content: space-between;
  gap: 20px;
  color: #a9bec9;
  font-size: 0.74rem;
  line-height: 1.5;
}

.anatomy-hero-media figcaption strong {
  font-size: 0.98rem;
}

.anatomy-index {
  position: relative;
  z-index: 3;
  display: grid;
  grid-template-columns: auto minmax(0, 1fr);
  align-items: center;
  gap: 24px;
  margin-bottom: -36px;
  padding: 15px 18px;
  border: 1px solid var(--anatomy-line);
  border-radius: 20px;
  background: rgba(5, 16, 26, 0.9);
  box-shadow: 0 20px 60px rgba(0, 0, 0, 0.24);
  backdrop-filter: blur(18px);
}

.anatomy-index-label {
  padding: 8px 11px;
  border-radius: 999px;
  background: rgba(207, 175, 112, 0.1);
  color: var(--anatomy-gold);
  font-size: 0.64rem;
  font-weight: 800;
  letter-spacing: 0.12em;
  text-transform: uppercase;
  white-space: nowrap;
}

.anatomy-index-links {
  display: flex;
  align-items: center;
  gap: 2px;
  overflow-x: auto;
  padding-bottom: 2px;
  scrollbar-width: thin;
  scrollbar-color: rgba(20, 241, 196, 0.32) transparent;
}

.anatomy-index-links a {
  flex: 0 0 auto;
  padding: 10px 11px;
  border-radius: 10px;
  color: #9db4bf;
  font-size: 0.76rem;
  font-weight: 650;
  text-decoration: none;
  transition: color 180ms ease, background 180ms ease;
}

.anatomy-index-links a:hover {
  background: rgba(20, 241, 196, 0.07);
  color: var(--anatomy-ink);
}

.anatomy-section {
  padding-block: clamp(88px, 10vw, 142px);
  border-top: 1px solid var(--anatomy-line);
}

.anatomy-section-deep {
  background:
    linear-gradient(180deg, rgba(3, 11, 18, 0.74), rgba(4, 14, 23, 0.48)),
    radial-gradient(circle at 12% 40%, rgba(31, 162, 255, 0.045), transparent 28rem);
}

.anatomy-section-map {
  padding-top: clamp(120px, 13vw, 180px);
  background:
    radial-gradient(circle at 50% 45%, rgba(18, 72, 77, 0.2), transparent 35rem),
    #02070c;
}

.anatomy-heading {
  display: grid;
  grid-template-columns: minmax(0, 0.9fr) minmax(300px, 0.55fr);
  align-items: end;
  gap: clamp(30px, 6vw, 86px);
  margin-bottom: clamp(40px, 6vw, 68px);
}

.anatomy-heading h2,
.anatomy-feature-copy h2,
.molt-copy h2,
.faq-layout > header h2 {
  margin: 0;
  color: var(--anatomy-ink);
  font-size: clamp(2.7rem, 5.5vw, 5.8rem);
  font-weight: 630;
  letter-spacing: -0.055em;
  line-height: 0.96;
  text-wrap: balance;
}

.anatomy-heading > p,
.faq-layout > header > p:last-child {
  margin: 0;
  color: var(--anatomy-muted);
  line-height: 1.75;
}

.anatomy-heading-wide {
  grid-template-columns: minmax(0, 1.1fr) minmax(320px, 0.55fr);
}

.body-map-shell {
  display: grid;
  grid-template-columns: minmax(0, 1.14fr) minmax(370px, 0.86fr);
  min-height: 690px;
  overflow: hidden;
  border: 1px solid rgba(170, 215, 229, 0.2);
  border-radius: 32px;
  background: #000;
  box-shadow: 0 38px 110px rgba(0, 0, 0, 0.42);
}

.body-map-stage {
  position: relative;
  min-width: 0;
  display: grid;
  place-items: center;
  overflow: hidden;
  padding: 48px;
  background:
    radial-gradient(circle at 50% 46%, rgba(20, 241, 196, 0.07), transparent 29rem),
    linear-gradient(155deg, #010203, #03080c);
}

.body-map-stage::before,
.body-map-stage::after {
  position: absolute;
  inset: 9%;
  border: 1px solid rgba(255, 255, 255, 0.055);
  border-radius: 50%;
  pointer-events: none;
  content: "";
}

.body-map-stage::after {
  inset: 20%;
}

.schematic-label {
  position: absolute;
  z-index: 2;
  top: 24px;
  left: 28px;
  margin: 0;
  color: #718c98;
  font-size: 0.65rem;
  font-weight: 760;
  letter-spacing: 0.14em;
  text-transform: uppercase;
}

.body-map-svg {
  position: relative;
  z-index: 1;
  width: min(100%, 670px);
  height: auto;
  overflow: visible;
  filter: drop-shadow(0 0 20px rgba(255, 255, 255, 0.08));
}

.map-placeholder-base {
  pointer-events: none;
  transition: opacity 260ms ease, filter 260ms ease;
}

.body-map-svg .map-part {
  fill: #fff;
  stroke: none;
  cursor: pointer;
  transition: opacity 260ms ease, filter 260ms ease, fill 260ms ease, stroke 260ms ease;
}

.body-map-svg .map-overlay {
  opacity: 0;
}

.body-map-svg .map-part-legs {
  fill: #fff;
  stroke: none;
}

.body-map-svg .map-part-pedipalps {
  fill: #fff;
}

.body-map-svg .map-part-eyes {
  stroke: none;
}

.body-map-svg .map-part-eyes .eye-tubercle {
  fill: #fff;
}

.body-map-svg .map-part-eyes circle {
  fill: #020406;
}

.body-map-svg .chelicera-seam {
  fill: none;
  stroke: #020406;
  stroke-width: 2;
  stroke-linecap: round;
}

.body-map-stage:not([data-active-part="whole"]) .map-part {
  opacity: 0.13;
}

.body-map-stage[data-active-part="prosoma"] .map-part-prosoma,
.body-map-stage[data-active-part="abdomen"] .map-part-abdomen,
.body-map-stage[data-active-part="legs"] .map-part-legs,
.body-map-stage[data-active-part="pedipalps"] .map-part-pedipalps,
.body-map-stage[data-active-part="chelicerae"] .map-part-chelicerae,
.body-map-stage[data-active-part="eyes"] .map-part-eyes,
.body-map-stage[data-active-part="spinnerets"] .map-part-spinnerets {
  opacity: 1;
  filter: drop-shadow(0 0 16px rgba(20, 241, 196, 0.82));
}

.body-map-stage[data-active-part="eyes"] .map-part-eyes .eye-tubercle {
  fill: var(--anatomy-teal);
}

.body-map-pulse {
  position: absolute;
  bottom: 30px;
  left: 30px;
  width: 7px;
  height: 7px;
  border-radius: 50%;
  background: var(--anatomy-teal);
  box-shadow: 0 0 16px rgba(20, 241, 196, 0.9);
}

.body-map-panel {
  display: flex;
  flex-direction: column;
  border-left: 1px solid var(--anatomy-line);
  background: linear-gradient(165deg, rgba(8, 25, 37, 0.94), rgba(3, 11, 18, 0.99));
}

.body-map-controls {
  display: grid;
  grid-template-columns: repeat(2, minmax(0, 1fr));
  gap: 8px;
  padding: 26px;
  border-bottom: 1px solid var(--anatomy-line);
}

.body-map-control {
  min-height: 47px;
  padding: 0 14px;
  border: 1px solid var(--anatomy-line);
  border-radius: 12px;
  background: rgba(255, 255, 255, 0.02);
  color: #a5bac4;
  font-size: 0.76rem;
  font-weight: 690;
  text-align: left;
  cursor: pointer;
  transition: border-color 180ms ease, background 180ms ease, color 180ms ease;
}

.body-map-control:hover,
.body-map-control.is-active {
  border-color: rgba(20, 241, 196, 0.42);
  background: rgba(20, 241, 196, 0.09);
  color: #eefbfa;
}

.body-map-reading {
  flex: 1;
  display: flex;
  flex-direction: column;
  justify-content: center;
  padding: clamp(34px, 5vw, 64px);
}

.body-map-reading-kicker {
  margin: 0 0 18px;
  color: var(--anatomy-gold);
  font-size: 0.68rem;
  font-weight: 780;
  letter-spacing: 0.15em;
  text-transform: uppercase;
}

.body-map-reading h3 {
  margin: 0;
  color: var(--anatomy-ink);
  font-size: clamp(2rem, 3.6vw, 3.5rem);
  font-weight: 620;
  letter-spacing: -0.045em;
  line-height: 1;
}

.body-map-reading > p:not(.body-map-reading-kicker) {
  margin: 22px 0 0;
  color: var(--anatomy-muted);
  line-height: 1.78;
}

.body-map-deep-link {
  align-self: flex-start;
  margin-top: 28px;
  color: var(--anatomy-teal);
  font-size: 0.78rem;
  font-weight: 730;
  text-decoration: none;
}

.anatomy-feature {
  display: grid;
  grid-template-columns: minmax(0, 1.1fr) minmax(380px, 0.9fr);
  align-items: stretch;
  gap: clamp(24px, 5vw, 72px);
}

.anatomy-feature-reverse {
  grid-template-columns: minmax(380px, 0.9fr) minmax(0, 1.1fr);
}

.anatomy-feature-copy {
  align-self: center;
}

.anatomy-feature-copy > p:not(.eyebrow),
.molt-copy > p:not(.eyebrow) {
  margin: 24px 0 0;
  color: var(--anatomy-muted);
  line-height: 1.78;
}

.anatomy-photo-frame {
  position: relative;
  overflow: hidden;
  margin: 0;
  border: 1px solid var(--anatomy-line);
  border-radius: 26px;
  background: #02070c;
  box-shadow: 0 30px 80px rgba(0, 0, 0, 0.26);
}

.anatomy-photo-frame img {
  width: 100%;
  height: 100%;
  display: block;
  object-fit: cover;
}

.anatomy-photo-frame figcaption {
  position: absolute;
  right: 14px;
  bottom: 14px;
  left: 14px;
  padding: 12px 14px;
  border: 1px solid rgba(255, 255, 255, 0.12);
  border-radius: 12px;
  background: rgba(2, 8, 13, 0.76);
  color: #a8bdc7;
  font-size: 0.69rem;
  line-height: 1.5;
  backdrop-filter: blur(12px);
}

.anatomy-photo-frame-wide {
  min-height: 560px;
}

.anatomy-photo-frame-portrait {
  min-height: 620px;
}

.anatomy-caution {
  padding: 18px 20px;
  border-left: 3px solid var(--anatomy-teal);
  border-radius: 0 12px 12px 0;
  background: rgba(20, 241, 196, 0.055);
  color: #a9bec7 !important;
  font-size: 0.86rem;
}

.anatomy-triptych {
  display: grid;
  grid-template-columns: repeat(3, minmax(0, 1fr));
  gap: 16px;
}

.anatomy-topic-card {
  min-width: 0;
  overflow: hidden;
  border: 1px solid var(--anatomy-line);
  border-radius: 24px;
  background: linear-gradient(155deg, rgba(9, 28, 42, 0.9), rgba(4, 13, 21, 0.98));
}

.anatomy-topic-card figure {
  margin: 0;
  border-bottom: 1px solid var(--anatomy-line);
  background: #020609;
}

.anatomy-topic-card figure img {
  width: 100%;
  height: auto;
  aspect-ratio: 1 / 0.75;
  display: block;
  object-fit: cover;
}

.anatomy-topic-card figcaption {
  padding: 11px 15px;
  color: #718e9b;
  font-size: 0.66rem;
  line-height: 1.45;
}

.anatomy-topic-card > div {
  padding: clamp(24px, 3.5vw, 38px);
}

.anatomy-card-number {
  color: var(--anatomy-gold);
  font-size: 0.67rem;
  font-weight: 790;
  letter-spacing: 0.14em;
}

.anatomy-topic-card h3,
.leg-notes h3,
.sexual-grid h3,
.photo-gap-note h3,
.related-anatomy-grid h3,
.lineage-anatomy-card h3 {
  margin: 18px 0 0;
  color: var(--anatomy-ink);
  font-size: clamp(1.55rem, 2.7vw, 2.5rem);
  font-weight: 630;
  letter-spacing: -0.04em;
  line-height: 1.03;
}

.anatomy-topic-card p,
.leg-notes p,
.sexual-grid p,
.photo-gap-note p,
.related-anatomy-grid p,
.lineage-anatomy-card li {
  color: var(--anatomy-muted);
  line-height: 1.72;
}

.anatomy-topic-card p {
  margin: 17px 0 0;
}

.leg-lab {
  overflow: hidden;
  border: 1px solid var(--anatomy-line);
  border-radius: 28px;
  background: #02070b;
}

.leg-model {
  display: grid;
  grid-template-columns: minmax(0, 1fr) 330px;
  min-height: 520px;
  background: #000;
}

.leg-model-stage {
  position: relative;
  min-width: 0;
  display: grid;
  place-items: center;
  padding: 74px 36px 36px;
  background:
    radial-gradient(circle at 52% 45%, rgba(20, 241, 196, 0.09), transparent 28rem),
    linear-gradient(150deg, #020507, #000);
}

.leg-model-svg {
  width: 100%;
  height: auto;
  overflow: visible;
}

.leg-joints circle {
  fill: #303b41;
  stroke: #94a3aa;
  stroke-width: 5;
}

.leg-model-part {
  fill: url(#leg-segment-light);
  stroke: rgba(255, 255, 255, 0.74);
  stroke-width: 2;
  stroke-linejoin: round;
  cursor: pointer;
  transition: opacity 220ms ease, filter 220ms ease, stroke 220ms ease;
}

.leg-model-trochanter,
.leg-model-patella,
.leg-model-metatarsus {
  fill: url(#leg-segment-mid);
}

.leg-model:not([data-active-segment="whole"]) .leg-model-part {
  opacity: 0.09;
}

.leg-model:not([data-active-segment="whole"]) .leg-joints,
.leg-model:not([data-active-segment="whole"]) .leg-model-highlights,
.leg-model:not([data-active-segment="whole"]) .leg-claws {
  opacity: 0.08;
}

.leg-model[data-active-segment="coxa"] .leg-model-coxa,
.leg-model[data-active-segment="trochanter"] .leg-model-trochanter,
.leg-model[data-active-segment="femur"] .leg-model-femur,
.leg-model[data-active-segment="patella"] .leg-model-patella,
.leg-model[data-active-segment="tibia"] .leg-model-tibia,
.leg-model[data-active-segment="metatarsus"] .leg-model-metatarsus,
.leg-model[data-active-segment="tarsus"] .leg-model-tarsus {
  opacity: 1;
  fill: #fff;
  stroke: var(--anatomy-teal);
  stroke-width: 6;
  filter: drop-shadow(0 0 18px rgba(20, 241, 196, 0.9));
}

.leg-model[data-active-segment="tarsus"] .leg-claws {
  opacity: 1;
  stroke: var(--anatomy-teal);
  filter: drop-shadow(0 0 10px rgba(20, 241, 196, 0.8));
}

.leg-model-highlights {
  fill: none;
  stroke: rgba(255, 255, 255, 0.7);
  stroke-width: 5;
  stroke-linecap: round;
  pointer-events: none;
}

.leg-claws {
  fill: none;
  stroke: #dce6e9;
  stroke-width: 6;
  stroke-linecap: round;
}

.leg-model-numbers text {
  fill: #071015;
  font-size: 26px;
  font-weight: 850;
  text-anchor: middle;
  dominant-baseline: middle;
  pointer-events: none;
}

.leg-model-panel {
  display: flex;
  flex-direction: column;
  border-left: 1px solid var(--anatomy-line);
  background: linear-gradient(165deg, rgba(8, 25, 37, 0.95), rgba(3, 11, 18, 0.99));
}

.leg-model-controls {
  display: grid;
  grid-template-columns: repeat(2, minmax(0, 1fr));
  gap: 7px;
  padding: 18px;
  border-bottom: 1px solid var(--anatomy-line);
}

.leg-model-control {
  min-height: 44px;
  display: flex;
  align-items: center;
  gap: 8px;
  padding: 8px 10px;
  border: 1px solid var(--anatomy-line);
  border-radius: 10px;
  background: rgba(255, 255, 255, 0.02);
  color: #a9bcc5;
  font: inherit;
  font-size: 0.68rem;
  font-weight: 700;
  cursor: pointer;
}

.leg-model-control span {
  width: 21px;
  height: 21px;
  flex: 0 0 auto;
  display: grid;
  place-items: center;
  border: 1px solid rgba(20, 241, 196, 0.32);
  border-radius: 50%;
  color: var(--anatomy-teal);
  font-size: 0.57rem;
}

.leg-model-control:hover,
.leg-model-control.is-active {
  border-color: rgba(20, 241, 196, 0.45);
  background: rgba(20, 241, 196, 0.09);
  color: #f2fffd;
}

.leg-model-reading {
  flex: 1;
  display: flex;
  flex-direction: column;
  justify-content: center;
  padding: 28px;
}

.leg-model-reading > p:first-child {
  margin: 0 0 12px;
  color: var(--anatomy-gold);
  font-size: 0.64rem;
  font-weight: 790;
  letter-spacing: 0.13em;
  text-transform: uppercase;
}

.leg-model-reading h3 {
  margin: 0;
  color: var(--anatomy-ink);
  font-size: 1.8rem;
  font-weight: 630;
  letter-spacing: -0.04em;
  line-height: 1;
}

.leg-model-reading > p:last-child {
  margin: 16px 0 0;
  color: var(--anatomy-muted);
  font-size: 0.84rem;
  line-height: 1.66;
}

.leg-notes {
  display: grid;
  grid-template-columns: repeat(3, minmax(0, 1fr));
  border-top: 1px solid var(--anatomy-line);
}

.leg-notes article {
  padding: clamp(28px, 4vw, 44px);
}

.leg-notes article + article {
  border-left: 1px solid var(--anatomy-line);
}

.leg-notes article > span,
.related-anatomy-grid > a > span {
  color: var(--anatomy-gold);
  font-size: 0.65rem;
  font-weight: 790;
  letter-spacing: 0.14em;
  text-transform: uppercase;
}

.leg-notes h3 {
  font-size: 1.55rem;
}

.leg-notes p {
  margin: 15px 0 0;
  font-size: 0.88rem;
}

.sense-plate {
  position: relative;
  height: 190px;
  overflow: hidden;
  margin-top: 30px;
  border: 1px solid var(--anatomy-line);
  border-radius: 20px;
  background: #000;
}

.sense-plate::after {
  position: absolute;
  right: 0;
  bottom: 42px;
  left: 0;
  height: 2px;
  background: rgba(255, 255, 255, 0.35);
  content: "";
}

.sense-hairs {
  position: absolute;
  right: 0;
  bottom: 44px;
  left: 0;
  display: flex;
  align-items: end;
  justify-content: space-evenly;
  padding-inline: 44px;
}

.sense-hairs span {
  width: 3px;
  height: 88px;
  display: block;
  border-radius: 999px;
  background: #fff;
  transform: rotate(-4deg);
  transform-origin: bottom;
  animation: hair-sway 2.8s ease-in-out infinite;
}

.sense-hairs span:nth-child(2) { height: 116px; animation-delay: -0.5s; }
.sense-hairs span:nth-child(3) { height: 72px; animation-delay: -1.1s; }
.sense-hairs span:nth-child(4) { height: 102px; animation-delay: -1.7s; }
.sense-hairs span:nth-child(5) { height: 82px; animation-delay: -2.2s; }

@keyframes hair-sway {
  0%, 100% { transform: rotate(-3deg); }
  50% { transform: rotate(4deg); }
}

.sense-wave {
  position: absolute;
  top: 30px;
  left: 28px;
  display: flex;
  gap: 8px;
}

.sense-wave i {
  width: 24px;
  height: 70px;
  display: block;
  border: 2px solid var(--anatomy-teal);
  border-top-color: transparent;
  border-bottom-color: transparent;
  border-radius: 50%;
  opacity: 0.16;
  animation: wave-pulse 2.4s ease-out infinite;
}

.sense-wave i:nth-child(2) { animation-delay: 0.25s; }
.sense-wave i:nth-child(3) { animation-delay: 0.5s; }

@keyframes wave-pulse {
  0% { opacity: 0; transform: scale(0.65); }
  35% { opacity: 0.65; }
  100% { opacity: 0; transform: scale(1.2); }
}

.sense-plate small {
  position: absolute;
  right: 14px;
  bottom: 10px;
  color: #607d89;
  font-size: 0.6rem;
  letter-spacing: 0.08em;
  text-transform: uppercase;
}

.silk-story {
  display: grid;
  grid-template-columns: minmax(0, 1.15fr) minmax(320px, 0.85fr);
  grid-template-rows: repeat(2, minmax(280px, auto));
  gap: 16px;
}

.silk-photo-main {
  grid-row: 1 / 3;
  min-height: 700px;
}

.silk-photo-detail {
  min-height: 310px;
}

.silk-schematic {
  position: relative;
  overflow: hidden;
  padding: 50px 30px 30px;
  border: 1px solid var(--anatomy-line);
  border-radius: 26px;
  background: #000;
}

.silk-schematic svg {
  width: 100%;
  height: 230px;
  display: block;
}

.silk-abdomen,
.silk-spinneret {
  fill: #fff;
}

.silk-thread {
  fill: none;
  stroke: var(--anatomy-teal);
  stroke-width: 3;
  stroke-linecap: round;
  stroke-dasharray: 280;
  animation: silk-draw 3.5s ease-in-out infinite;
}

.silk-thread-2,
.silk-thread-4 { animation-delay: -1.1s; }

@keyframes silk-draw {
  0% { opacity: 0; stroke-dashoffset: 280; }
  35%, 80% { opacity: 0.9; stroke-dashoffset: 0; }
  100% { opacity: 0; stroke-dashoffset: -60; }
}

.silk-schematic > p:last-child {
  margin: 12px 0 0;
  color: var(--anatomy-muted);
  font-size: 0.84rem;
  line-height: 1.68;
}

.lung-schematic {
  position: relative;
  margin-top: 32px;
  padding: 58px 20px 20px;
  overflow: hidden;
  border: 1px solid var(--anatomy-line);
  border-radius: 20px;
  background: #000;
}

.lung-schematic svg {
  width: 100%;
  height: auto;
  display: block;
}

.lung-chamber {
  fill: rgba(255, 255, 255, 0.025);
  stroke: rgba(255, 255, 255, 0.42);
  stroke-width: 3;
}

.lung-opening {
  fill: none;
  stroke: #fff;
  stroke-width: 9;
  stroke-linecap: round;
}

.lung-route {
  fill: none;
  stroke: var(--anatomy-teal);
  stroke-width: 5;
  stroke-dasharray: 12 10;
  animation: lung-airflow 1.8s linear infinite;
}

@keyframes lung-airflow {
  to { stroke-dashoffset: -44; }
}

.lung-lamellae path {
  fill: none;
  stroke: #edf4f5;
  stroke-width: 8;
  stroke-linecap: round;
  filter: drop-shadow(0 0 5px rgba(255, 255, 255, 0.12));
}

.lung-air-particles circle {
  fill: var(--anatomy-teal);
  filter: drop-shadow(0 0 5px rgba(20, 241, 196, 0.8));
  animation: lung-particle 2.4s linear infinite;
}

.lung-air-particles circle:nth-child(2) { animation-delay: -0.6s; }
.lung-air-particles circle:nth-child(3) { animation-delay: -1.2s; }
.lung-air-particles circle:nth-child(4) { animation-delay: -1.8s; }

@keyframes lung-particle {
  0% { opacity: 0; transform: translateX(-25px); }
  25%, 75% { opacity: 1; }
  100% { opacity: 0; transform: translateX(75px); }
}

.lung-exchange {
  fill: none;
  stroke-width: 4;
  stroke-dasharray: 7 6;
}

.lung-exchange-o2 { stroke: #78b5f0; }
.lung-exchange-co2 { stroke: #f293d1; }

.lung-svg-label {
  fill: #8ca6b2;
  font-size: 17px;
  font-weight: 720;
  letter-spacing: 0.02em;
}

.lung-gas-label {
  font-size: 22px;
  font-weight: 820;
}

.lung-gas-o2 { fill: #78b5f0; }
.lung-gas-co2 { fill: #f293d1; }

.lung-explanation {
  display: grid;
  grid-template-columns: repeat(3, minmax(0, 1fr));
  gap: 8px;
  margin: 8px 0 0;
}

.lung-explanation span {
  display: flex;
  align-items: flex-start;
  gap: 8px;
  color: #819ca8;
  font-size: 0.67rem;
  line-height: 1.45;
}

.lung-explanation b {
  width: 20px;
  height: 20px;
  flex: 0 0 auto;
  display: grid;
  place-items: center;
  border: 1px solid rgba(20, 241, 196, 0.4);
  border-radius: 50%;
  color: var(--anatomy-teal);
  font-size: 0.57rem;
}

.internal-map {
  display: grid;
  grid-template-columns: minmax(0, 1fr) minmax(360px, 0.82fr);
  overflow: hidden;
  border: 1px solid var(--anatomy-line);
  border-radius: 28px;
  background: #000;
}

.internal-figure {
  position: relative;
  min-width: 0;
  display: grid;
  place-items: center;
  padding: 56px 4px 4px;
  background: radial-gradient(circle at 55% 48%, rgba(20, 241, 196, 0.06), transparent 27rem);
}

.internal-figure svg {
  width: min(112%, 760px);
  max-width: none;
  height: auto;
  display: block;
}

.internal-vector-base {
  fill: #fff;
  opacity: 0.24;
  pointer-events: none;
  transition: opacity 220ms ease, filter 220ms ease;
}

.internal-map:not([data-active-system="whole"]) .internal-vector-base {
  opacity: 0.14;
}

.internal-region-guide path {
  fill: none;
  stroke: rgba(255, 255, 255, 0.44);
  stroke-width: 1.5;
  stroke-dasharray: 5 5;
}

.internal-region-guide text {
  fill: #86a3af;
  font-size: 10px;
  font-weight: 780;
  letter-spacing: 0.12em;
}

.internal-spider-legs {
  fill: none;
  stroke: rgba(255, 255, 255, 0.29);
  stroke-width: 17;
  stroke-linecap: square;
  stroke-linejoin: miter;
}

.internal-outline {
  fill: rgba(255, 255, 255, 0.035);
  stroke: rgba(255, 255, 255, 0.72);
  stroke-width: 5;
}

.internal-pedicel {
  fill: rgba(255, 255, 255, 0.08);
  stroke: rgba(255, 255, 255, 0.72);
  stroke-width: 4;
}

.internal-spinnerets {
  fill: rgba(255, 255, 255, 0.05);
  stroke: rgba(255, 255, 255, 0.72);
  stroke-width: 4;
}

.internal-region-label {
  fill: rgba(255, 255, 255, 0.45);
  font-size: 15px;
  font-weight: 780;
  letter-spacing: 0.14em;
  text-anchor: middle;
}

.internal-pedicel-label {
  fill: #7e98a4;
  font-size: 13px;
  font-weight: 680;
}

.internal-system {
  fill: none;
  stroke-width: 8;
  stroke-linecap: round;
  stroke-linejoin: round;
  cursor: pointer;
  transition: opacity 220ms ease, filter 220ms ease;
}

.internal-system-nerves { --system-color: #14f1c4; fill: rgba(20, 241, 196, 0.16); stroke: #14f1c4; }
.internal-system-venom { --system-color: #d6af5f; fill: rgba(214, 175, 95, 0.16); stroke: #d6af5f; }
.internal-system-heart { --system-color: #f56974; fill: rgba(245, 105, 116, 0.18); stroke: #f56974; }
.internal-system-gut { --system-color: #f1f6f8; stroke: #f1f6f8; }
.internal-system-lungs { --system-color: #78b5f0; fill: rgba(120, 181, 240, 0.17); stroke: #78b5f0; }
.internal-system-silk { --system-color: #ad84ef; fill: rgba(173, 132, 239, 0.16); stroke: #ad84ef; }
.internal-system-reproduction { --system-color: #f293d1; fill: rgba(242, 147, 209, 0.16); stroke: #f293d1; }

.internal-system-lungs {
  stroke-dasharray: 7 6;
}

.internal-map:not([data-active-system="whole"]) .internal-system {
  opacity: 0.06;
}

.internal-map[data-active-system="nerves"] .internal-system-nerves,
.internal-map[data-active-system="venom"] .internal-system-venom,
.internal-map[data-active-system="heart"] .internal-system-heart,
.internal-map[data-active-system="gut"] .internal-system-gut,
.internal-map[data-active-system="lungs"] .internal-system-lungs,
.internal-map[data-active-system="silk"] .internal-system-silk,
.internal-map[data-active-system="reproduction"] .internal-system-reproduction {
  opacity: 1;
  stroke-width: 14;
  filter: drop-shadow(0 0 16px var(--system-color));
}

.internal-map[data-active-system="nerves"] .internal-system-nerves { fill: rgba(20, 241, 196, 0.42); }
.internal-map[data-active-system="venom"] .internal-system-venom { fill: rgba(214, 175, 95, 0.48); }
.internal-map[data-active-system="heart"] .internal-system-heart { fill: rgba(245, 105, 116, 0.55); }
.internal-map[data-active-system="lungs"] .internal-system-lungs { fill: rgba(120, 181, 240, 0.58); }
.internal-map[data-active-system="silk"] .internal-system-silk { fill: rgba(173, 132, 239, 0.45); }
.internal-map[data-active-system="reproduction"] .internal-system-reproduction { fill: rgba(242, 147, 209, 0.42); }

.internal-key {
  border-left: 1px solid var(--anatomy-line);
  background: linear-gradient(165deg, rgba(8, 25, 37, 0.94), rgba(3, 11, 18, 0.99));
}

.internal-key-item {
  width: 100%;
  display: block;
  padding: 18px 22px;
  border-top: 1px solid var(--anatomy-line);
  border-right: 0;
  border-bottom: 0;
  border-left: 0;
  background: transparent;
  color: inherit;
  font: inherit;
  text-align: left;
  cursor: pointer;
  transition: background 180ms ease;
}

.internal-key-item:first-child {
  border-top: 0;
}

.internal-key-item:hover,
.internal-key-item.is-active {
  background: rgba(20, 241, 196, 0.075);
}

.internal-key-title {
  display: flex;
  align-items: center;
  gap: 10px;
  color: var(--anatomy-ink);
  font-size: 0.86rem;
  font-weight: 730;
}

.internal-key-title i {
  width: 9px;
  height: 9px;
  flex: 0 0 auto;
  display: block;
  border-radius: 50%;
}

.internal-key-title small {
  margin-left: auto;
  color: #6f8b98;
  font-size: 0.58rem;
  font-weight: 760;
  letter-spacing: 0.08em;
  text-transform: uppercase;
}

.key-whole { background: #8ca5b1; }
.key-nerves { background: #14f1c4; }
.key-venom { background: #d6af5f; }
.key-heart { background: #f56974; }
.key-gut { background: #f1f6f8; }
.key-lungs { background: #78b5f0; }
.key-silk { background: #ad84ef; }
.key-reproduction { background: #f293d1; }

.internal-key-copy {
  display: block;
  margin: 7px 0 0 19px;
  color: #8ca5b1;
  font-size: 0.79rem;
  line-height: 1.58;
}

.molt-layout {
  display: grid;
  grid-template-columns: minmax(0, 1.05fr) minmax(380px, 0.95fr);
  align-items: stretch;
  gap: clamp(26px, 5vw, 72px);
}

.molt-photo {
  min-height: 760px;
}

.molt-copy {
  align-self: center;
}

.molt-steps {
  display: grid;
  gap: 0;
  margin: 32px 0 0;
  padding: 0;
  list-style: none;
}

.molt-steps li {
  display: grid;
  grid-template-columns: auto minmax(0, 1fr);
  gap: 20px;
  padding: 18px 0;
  border-top: 1px solid var(--anatomy-line);
}

.molt-steps li > span {
  color: var(--anatomy-gold);
  font-size: 0.67rem;
  font-weight: 790;
  letter-spacing: 0.13em;
}

.molt-steps strong {
  display: block;
  font-size: 0.88rem;
}

.molt-steps p {
  margin: 6px 0 0;
  color: var(--anatomy-muted);
  font-size: 0.82rem;
  line-height: 1.55;
}

.lineage-anatomy-grid {
  display: grid;
  grid-template-columns: repeat(2, minmax(0, 1fr));
  gap: 16px;
}

.lineage-anatomy-card {
  overflow: hidden;
  border: 1px solid var(--anatomy-line);
  border-radius: 25px;
  background: linear-gradient(155deg, rgba(9, 28, 42, 0.9), rgba(4, 13, 21, 0.98));
}

.lineage-anatomy-card figure {
  margin: 0;
  border-bottom: 1px solid var(--anatomy-line);
  background: #000;
}

.lineage-anatomy-card figure img {
  width: 100%;
  height: auto;
  aspect-ratio: 16 / 9;
  display: block;
  object-fit: cover;
}

.lineage-anatomy-card > div {
  padding: clamp(28px, 4vw, 46px);
}

.lineage-anatomy-card h3 {
  margin-top: 14px;
}

.lineage-anatomy-card ul {
  display: grid;
  gap: 12px;
  margin: 24px 0 0;
  padding-left: 1.15rem;
}

.lineage-anatomy-card li {
  padding-left: 5px;
}

.lineage-comparison-plate {
  max-width: 520px;
  overflow: hidden;
  margin: 18px auto 0;
  border: 1px solid var(--anatomy-line);
  border-radius: 22px;
  background: #000;
}

.lineage-comparison-plate img {
  width: 100%;
  height: auto;
  display: block;
}

.lineage-comparison-plate figcaption {
  padding: 14px 17px;
  border-top: 1px solid var(--anatomy-line);
  color: #77929f;
  font-size: 0.68rem;
  line-height: 1.5;
}

.lineage-caution {
  max-width: 900px;
  margin: 28px auto 0;
}

.sexual-grid {
  display: grid;
  grid-template-columns: repeat(3, minmax(0, 1fr));
  gap: 16px;
}

.sexual-grid article {
  min-height: 340px;
  padding: clamp(28px, 4vw, 44px);
  border: 1px solid var(--anatomy-line);
  border-radius: 23px;
  background: linear-gradient(155deg, rgba(9, 28, 42, 0.9), rgba(4, 13, 21, 0.98));
}

.sexual-mark {
  width: 48px;
  height: 48px;
  display: grid;
  place-items: center;
  border: 1px solid rgba(20, 241, 196, 0.34);
  border-radius: 50%;
  color: var(--anatomy-teal);
  font-size: 1.35rem;
  box-shadow: 0 0 24px rgba(20, 241, 196, 0.08);
}

.sexual-mark-neutral {
  border-color: rgba(207, 175, 112, 0.34);
  color: var(--anatomy-gold);
}

.sexual-grid p {
  margin: 20px 0 0;
}

.photo-gap-note {
  display: grid;
  grid-template-columns: minmax(180px, 0.35fr) minmax(240px, 0.65fr) minmax(0, 1fr);
  align-items: center;
  gap: 30px;
  margin-top: 18px;
  padding: clamp(26px, 4vw, 42px);
  border: 1px solid rgba(207, 175, 112, 0.24);
  border-radius: 22px;
  background: rgba(207, 175, 112, 0.04);
}

.photo-gap-note .eyebrow,
.photo-gap-note h3,
.photo-gap-note p {
  margin: 0;
}

.photo-gap-note h3 {
  font-size: clamp(1.45rem, 2.5vw, 2.15rem);
}

.faq-layout {
  display: grid;
  grid-template-columns: minmax(300px, 0.7fr) minmax(0, 1.3fr);
  align-items: start;
  gap: clamp(36px, 7vw, 100px);
}

.faq-layout > header {
  position: sticky;
  top: 116px;
}

.faq-layout > header > p:last-child {
  margin-top: 24px;
}

.anatomy-faq {
  border-top: 1px solid var(--anatomy-line);
}

.anatomy-faq details {
  border-bottom: 1px solid var(--anatomy-line);
}

.anatomy-faq summary {
  position: relative;
  padding: 25px 50px 25px 0;
  color: var(--anatomy-ink);
  font-size: clamp(1rem, 1.7vw, 1.2rem);
  font-weight: 690;
  cursor: pointer;
  list-style: none;
}

.anatomy-faq summary::-webkit-details-marker {
  display: none;
}

.anatomy-faq summary::after {
  position: absolute;
  top: 50%;
  right: 5px;
  color: var(--anatomy-teal);
  font-size: 1.35rem;
  font-weight: 350;
  content: "+";
  transform: translateY(-50%);
}

.anatomy-faq details[open] summary::after {
  content: "−";
}

.anatomy-faq p {
  margin: -5px 60px 26px 0;
  color: var(--anatomy-muted);
  line-height: 1.7;
}

.anatomy-section-related {
  background: radial-gradient(circle at 50% 50%, rgba(20, 241, 196, 0.05), transparent 34rem);
}

.related-anatomy-grid {
  display: grid;
  grid-template-columns: repeat(3, minmax(0, 1fr));
  gap: 16px;
}

.related-anatomy-grid > a {
  min-height: 310px;
  display: flex;
  flex-direction: column;
  padding: clamp(28px, 4vw, 42px);
  border: 1px solid var(--anatomy-line);
  border-radius: 22px;
  background: linear-gradient(155deg, rgba(9, 28, 42, 0.86), rgba(4, 13, 21, 0.96));
  color: inherit;
  text-decoration: none;
  transition: transform 180ms ease, border-color 180ms ease, background 180ms ease;
}

.related-anatomy-grid > a:hover {
  transform: translateY(-4px);
  border-color: rgba(20, 241, 196, 0.36);
  background: linear-gradient(155deg, rgba(11, 37, 49, 0.92), rgba(4, 15, 23, 0.99));
}

.related-anatomy-grid h3 {
  margin-top: 32px;
}

.related-anatomy-grid p {
  margin: 17px 0 0;
}

.related-anatomy-grid strong {
  margin-top: auto;
  padding-top: 28px;
  color: var(--anatomy-teal);
  font-size: 0.76rem;
}

.anatomy-sources > p:not(.eyebrow) {
  max-width: 780px;
}

@media (max-width: 1060px) {
  .anatomy-hero {
    grid-template-columns: minmax(0, 0.78fr) minmax(450px, 1.22fr);
  }

  .body-map-shell {
    grid-template-columns: minmax(0, 1fr) 350px;
  }

  .anatomy-triptych,
  .sexual-grid,
  .related-anatomy-grid {
    grid-template-columns: repeat(2, minmax(0, 1fr));
  }

  .anatomy-topic-card:last-child,
  .sexual-grid article:last-child,
  .related-anatomy-grid > a:last-child {
    grid-column: 1 / -1;
  }
}

@media (max-width: 900px) {
  .anatomy-hero,
  .anatomy-feature,
  .anatomy-feature-reverse,
  .molt-layout,
  .faq-layout {
    grid-template-columns: 1fr;
  }

  .anatomy-hero {
    min-height: auto;
  }

  .anatomy-hero-media {
    min-height: 620px;
  }

  .anatomy-index {
    grid-template-columns: 1fr;
    gap: 6px;
  }

  .anatomy-index-label {
    justify-self: start;
  }

  .anatomy-heading,
  .anatomy-heading-wide {
    grid-template-columns: 1fr;
    align-items: start;
  }

  .body-map-shell {
    grid-template-columns: 1fr;
  }

  .body-map-stage {
    min-height: 650px;
  }

  .body-map-panel {
    border-top: 1px solid var(--anatomy-line);
    border-left: 0;
  }

  .body-map-reading {
    min-height: 340px;
  }

  .anatomy-feature-reverse .anatomy-feature-copy {
    order: 2;
  }

  .anatomy-photo-frame-wide,
  .anatomy-photo-frame-portrait,
  .molt-photo {
    min-height: 600px;
  }

  .silk-story {
    grid-template-columns: 1fr;
    grid-template-rows: auto;
  }

  .silk-photo-main {
    grid-row: auto;
    min-height: 620px;
  }

  .silk-photo-detail {
    min-height: 430px;
  }

  .internal-map {
    grid-template-columns: 1fr;
  }

  .internal-key {
    display: grid;
    grid-template-columns: repeat(2, minmax(0, 1fr));
    border-top: 1px solid var(--anatomy-line);
    border-left: 0;
  }

  .internal-key-item:nth-child(2) {
    border-top: 0;
  }

  .internal-key-item:nth-child(even) {
    border-left: 1px solid var(--anatomy-line);
  }

  .faq-layout > header {
    position: static;
  }

  .photo-gap-note {
    grid-template-columns: 1fr;
    gap: 14px;
  }
}

@media (max-width: 680px) {
  .anatomy-hero {
    padding-block: 62px 88px;
  }

  .anatomy-hero-copy h1 {
    font-size: clamp(3.5rem, 18vw, 5.7rem);
  }

  .anatomy-hero-media {
    min-height: 520px;
    border-radius: 21px;
  }

  .anatomy-hero-media figcaption {
    align-items: start;
    flex-direction: column;
    gap: 6px;
  }

  .anatomy-index {
    width: calc(100% - 28px);
    margin-bottom: -26px;
    border-radius: 16px;
  }

  .anatomy-section {
    padding-block: 80px;
  }

  .anatomy-section-map {
    padding-top: 112px;
  }

  .anatomy-heading h2,
  .anatomy-feature-copy h2,
  .molt-copy h2,
  .faq-layout > header h2 {
    font-size: clamp(2.7rem, 14vw, 4.5rem);
  }

  .body-map-shell,
  .leg-lab,
  .internal-map {
    border-radius: 20px;
  }

  .body-map-stage {
    min-height: 470px;
    padding: 52px 22px 24px;
  }

  .schematic-label {
    top: 18px;
    left: 20px;
  }

  .body-map-controls {
    padding: 16px;
  }

  .body-map-reading {
    min-height: 330px;
    padding: 32px 24px;
  }

  .anatomy-triptych,
  .sexual-grid,
  .related-anatomy-grid,
  .lineage-anatomy-grid {
    grid-template-columns: 1fr;
  }

  .anatomy-topic-card:last-child,
  .sexual-grid article:last-child,
  .related-anatomy-grid > a:last-child {
    grid-column: auto;
  }

  .anatomy-photo-frame {
    border-radius: 20px;
  }

  .anatomy-photo-frame-wide,
  .anatomy-photo-frame-portrait,
  .molt-photo,
  .silk-photo-main,
  .silk-photo-detail {
    min-height: 450px;
  }

  .leg-model {
    grid-template-columns: 1fr;
  }

  .leg-model-stage {
    min-height: 360px;
    padding: 70px 16px 22px;
  }

  .leg-model-panel {
    border-top: 1px solid var(--anatomy-line);
    border-left: 0;
  }

  .leg-model-controls {
    grid-template-columns: repeat(2, minmax(0, 1fr));
    padding: 14px;
  }

  .leg-model-reading {
    min-height: 240px;
    padding: 28px 22px;
  }

  .leg-notes {
    grid-template-columns: 1fr;
  }

  .leg-notes article + article {
    border-top: 1px solid var(--anatomy-line);
    border-left: 0;
  }

  .lung-schematic {
    padding: 54px 10px 18px;
  }

  .lung-explanation {
    grid-template-columns: 1fr;
    gap: 10px;
    padding-inline: 10px;
  }

  .internal-figure {
    min-height: 390px;
    padding: 58px 12px 20px;
  }

  .internal-key {
    grid-template-columns: 1fr;
  }

  .internal-key-item:nth-child(2) {
    border-top: 1px solid var(--anatomy-line);
  }

  .internal-key-item:nth-child(even) {
    border-left: 0;
  }

  .internal-key-title small {
    display: none;
  }

  .lineage-comparison-plate {
    max-width: 100%;
  }

  .anatomy-faq p {
    margin-right: 0;
  }
}

@media (prefers-reduced-motion: reduce) {
  .body-map-svg .map-part,
  .leg-model-part,
  .sense-hairs span,
  .sense-wave i,
  .silk-thread,
  .lung-route,
  .lung-air-particles circle,
  .internal-system,
  .related-anatomy-grid > a {
    animation: none;
    transition: none;
  }

  .silk-thread {
    opacity: 0.85;
    stroke-dashoffset: 0;
  }
}